"Hence, late-surviving populations of H. heidelbergensis/H. rhodesiensis could have been a source of 'ghost' introgressions" the authors write.
I have repeatedly argued that.
I have continually opined that
Homo Habilis and
Homo Erectus were alive as late as 12,000 years ago.
There is a unique language family in Africa known as click-languages.
The Xhosa tribe in south Africa and one or two other tribes near the Kalahari Desert are the only ones that use them.
Neither
Homo Habilis nor
Homo Erectus could speak, because they lacked a hyoid bone and the FOXP2 gene, but that does not mean they could not communicate.
They could communicate by making clicking noises using their tongues.
I would suggest they did exactly that, and that bands of Xhosa and others were in contact with bands or family groups of
Homo Habilis or
Homo Erectus and that they learned to communicate with
Homo Habilis and
Homo Erectus using the clicking noises, then later incorporated the clicking sounds into their own languages.
I also argue hominids emerged independently in Africa and Asia (Southeast Asia and Central Asia).